HALIFAX -- An investigation has found that the deaths of salmon and trout at three Nova Scotia aquaculture sites earlier this year were caused by environmental factors, not disease.

Staff at the provincial Fisheries Department say the fish kill was caused by extremely cold temperatures experienced during a particularly harsh winter.

Fisheries Minister Keith Colwell says no significant disease agents were found at any of the sites.

The sites, operated by New Brunswick's Cooke Aquaculture, are at Shelburne Harbour, Jordan Bay and Port Wade.

The company has said it's unclear how many fish died, though it appears the majority of the fish at the three sites survived.

Colwell says financial losses from the fish kill won't be covered by the province.